The Forest
Redwood forests generate the world’s greatest volume of living
matter per unit of land surface.
Redwoods tend to dominate their environment because of their resistance
to fire, drought, shade and disease and their ability to reproduce
from burls. They will survive all but the very worst floods or fires
and they can live with very little sunlight for decades. On floodplains
they will often grow almost exclusively. In areas where they grow
less vigorously such as on steeper slopes or after logging, many
other plant species have learned to live alongside the redwoods.
Douglas firs are one of the most common and may rival the height
of the redwoods in some drier areas. The Broad-leaf tan Oak is the
most successful tree under the canopy - it grows very well in the
shade and will grow in the decaying leaf matter, not tolerated by
most other plants. Other trees include the Western hemlock, California
laurel, maple, alder, huckleberry, Sitka spruce and rhododendron.
The forest floor is covered in ferns, evergreen huckleberry, salal,
and the western azalea. Wildflowers include redwood violets, trilliums,
Douglas iris, calypso orchids, and redwood sorrel.
The floor is covered in ‘duff’ – the decaying
leaves and twigs. This duff absorbs much of the water and keeps
roots moist as well as protecting them from erosion. This layer
contains many fungi and bacteria, which convert the dead plant and
animal matter into nutrients. A fallen redwood tree may survive
for hundreds of years on the forest floor.
Reproduction
One of the reasons the Coast Redwood has survived is its ability
to reproduce in two different ways: seed or sprout
Seed Reproduction:
This is the primary means of reproduction in an old-growth forest,
and it allows the genetic variation which is necessary for the long
term survival of the species. If the tree is exposed to enough sunlight
it can produce seeds at around 20 years of age. Seed production
peaks at 250 - 1000 years. Redwoods have both male (pollen) and
female (ovulate) cones, which grow on separate branches of the same
tree. They form on the tips of high branches on mature redwoods
in the fall. Pollen from the peppercorn-sized male cones is released
in streams in the fall. It drifts and descends through the canopy
to settle in the female cones. Fertilization will happen 4-8 weeks
later.
Over the next year, the female cones mature into hard-sided cones
from .5 to 1.5 inches in length. Protected by the hard scales are
anywhere from 60 to 150 seeds. The seeds of the redwood are tiny,
about the size of a small oatmeal flake. The will remain viable
for up to a year. Some seeds are released from the cone while the
cone is still on the tree, drifting down to the ground or blown
by the wind. Most will settle within a few hundred feet of the tree.
Other seeds remain until the cone falls to the forest floor, dries,
and opens.
Less than 1% of these seeds will ever produce a tree. Many seeds
cannot penetrate the leaf litter or duff to reach the soil. Some
will land in water or on animals and will be carried away from the
redwood forest. Many more will reach the soil but will be destroyed
by fungal attack, bacteria, and insects. Many will be washed away
by rain, and some will be eaten by animals. Probably fewer than
one in a million seeds will ever become mature trees.
Seedlings will grow best under shady conditions, under the protection
of larger trees. When larger trees fall, opening light gaps, seedlings
will spring up to fill the void. They can grow roots of over an
inch after just one day. Most seedlings will die within 3 months.
Lack of moisture is the most likely cause, although many seedlings
will also be washed away, eaten by animals or attacked by fungi.
Their best chance of survival is after a fire, flood or fallen tree,
when most of the competition is gone.
After a year the roots will be around 1 ft deep and the stem around
5 inches. After their first few years, redwoods can grow very rapidly,
2 to 3 feet in height and up to one inch in diameter a year. The
trees can be 30ft (10m) at 20 years, 100ft (30m) at 50 years and
fully grown at 200-300. After 300 years the tree will grow wider
but not taller.
Sprout (burl) Reproduction:
Redwoods are one of a very few coniferous trees that can also reproduce
by sprouting. As early as one year after beginning to grow, a redwood
can begin to produce bud collars, most commonly referred to as ‘burls’.
These dormant root buds will continuously form, sometimes creating
large bulbous growth on the tree's roots, base, or trunk. Hormones
within the tree keep these burls from sprouting until the tree faces
some form of stress. Fire, erosion, flooding, browsing, or other
injury to the tree will trigger a release on the buds, causing them
to begin to grow within a few weeks.
Redwood sprouts, or suckers, can grow rapidly, receiving their early
nutrition from the roots on which they grow. They are genetically
identical to their parent tree, often forming "family circles"
around the parent tree. Under optimal conditions, sprouts may reach
heights of 8-10 feet in their first year. Eventually the tree will
grow its own root system and can survive even if the parent tree
should topple. After a fire trees will often sprout in such proliferation
that many of the sprouts cannot compete and die back. Some day these
sprouts will grow their own burls and produce their own sprouts.
Most sprouting will occur from a tree 200 – 400 years old.
Fire
Frequent fires ignited by lightning are an essential part of the
forest ecosystem. These fires will remove less fire resistant trees,
recycle nutrients, and modify the ground vegetation. The 12 inch
thick bark will protect the tree from all but the most intense fires.
The foliage is protected from fire by its distance from the forest
floor. Major fires occur anywhere from every 100 to 500 years depending
on the local conditions. The last significant fire in the Muir Woods
was around 1845. Most damage is done to seedlings and young trees,
which lack the defense of the mature trees.
It is a matter of debate whether the forest requires these fires
to survive (Unlike the Giant Sequoias which cannot reproduce without
fire). After a fire the redwood seedlings will usually be the first
to grow. They will take advantage of the reduced competition for
moisture and sunlight.
Fire ecology has changed throughout the years. It used to be assumed
that all fires were bad – they were either prevented or extinguished
as soon as possible. However, in the last few decades, the value
of fire has been recognized, as it burns up the fuel, reducing the
chance of a canopy fire and it returns many important nutrients
to the soil. Today, there is a fuel break surrounding the National
Monument. This break stops fires from spreading into Muir Woods
from the surrounding Mt Tamalpais. The break is maintained through
controlled fires, which are set in cool weather the burn the ground
debris.
Flood
The redwoods are also excellent survivors of the frequent floods
through the valleys. The floodwater will clear out much of the groundcover,
leaving new alluvial deposits, rich in nutrients for the new seedlings
to thrive. A severe flood can often bring up to 4ft of new sediment,
which will suffocate the roots of most trees. Redwoods have the
almost unique ability to grow a new layer of roots from the trunk
to penetrate the new soil.
Logging
The Coast Redwood has long been a popular target for the lumber
companies. The trees along with the Douglas fir are the most important
in the state. Redwood lumber is very popular due to its rich color,
durability and resistance to termites and decay. Partly because
of it commercial value, lumber companies have done a large amount
research into the redwood. Most of today’s redwoods are now
protected from logging but only 5% of the original old growth redwoods
exist today.
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